This paper has been reviewed by two reviewers. As Subject Editor handling the paper, your task is now to review the reviewers' comments and add comments of your own as necessary. These may be issues with the text that you pick up that the editors have not commented on, or responses from you to the author based on the reviewer comments (for example, agreeing, disagreeing, or clarifying the reviewer comments).

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The reviewers' recommendations are given at the bottom of the page. Please also add you own general comments to the author, as necessary.

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Remember, you need to assess each taxon (if any) as well as the main body of the paper.

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When done, check the 'Done' checkbox, and Submit.